Harmful blooms of the filamentous cyanobacteria Planktothrix rubescens have become common in many lakes as they have recovered from eutrophication over the last decades. These cyanobacteria, capable of regulating their vertical position, often flourish at the thermocline to form a deep chlorophyll maximum. In Lake Zurich (Switzerland), they accumulate during stratified season (May–October) as a persistent metalimnetic thin lay...