The consequences of hydrodynamic effects on survival of embryonic benthic organisms remain unknown. This is because the interaction of hydrodynamics and substrate roughness is complex, but it can be conceptualized in two dimensions by roughness flow regimes (isolated, wake interference, and skimming flow) or roughness types ( k ‐ and d ‐type). We examined the effect of different roughness flow regimes/roughness types and rough...