The isotopic and elemental composition of the fossil shells of foraminifera are often used for reconstructing past environments and climates. These so-called proxy relations are based on the effect of environmental conditions (e.g. seawater temperature, pH) on the isotopic ratio (e.g. δ 11 B or δ 18 O) or partitioning of elements (commonly expressed as El/Ca or D El ) during calcification. Whereas many studies focused on proxy...