Summary SR protein‐specific kinases (SRPKs) uniquely with a spacer region are important splicing factors from yeast to human. However, little is known about their biological functions in filamentous fungi. Therefore, we characterized a SRPK called SRK1 in wheat scab fungus Fusarium graminearum . Our data showed that Srk1 is required for vegetative growth, sexual reproduction and plant infection, and plays critical roles in pre...