Nitrogen and phosphorus budgets were compiled for the littoral (29 km 2 ) and pelagic (329 km 2 ) of ancient, deep, clear, and hard water Lake Ohrid (Albania and North Macedonia), to assess the importance of the littoral in nutrient retention. P originates mainly from domestic point sources (73%), for N this is karst seepage (50%). Total littoral loads are estimated at 1700 kg P and 23,200 kg N km −2 (area of littoral) yr −1 ;...