Narcomedusae play a key role as top‐down regulators in the midwater, the largest and most understudied biome on Earth. Here, we used ecological niche modeling in three‐dimensions (3D), ecomorphology, and phylogeny, to answer evolutionary and ecological questions about the widespread narcomedusan genus Solmissus . Our phylogenetic analyses confirmed that Solmissus incisa represents a complex of several cryptic species. Both the...