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The nitrogen isotopic composition of shell-bound organic matter in planktonic foraminifera (FB-δ 15 N) is widely used as a proxy for past ocean deoxygenation because water-column denitrification in oxygen-deficient zones (ODZs; [O 2 ] < 5 µmol/kg) preferentially removes 14 N, enriching the remaining nitrate in 15 N. Typically, increases in FB-δ 15 N records from ODZ-influenced regions are interpreted as evidence of ODZ expansi...
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