Seventeen Middle Pleistocene hominin teeth were excavated at Pontnewydd Cave in North Wales between 1980 and 1995 with a probable age of ∼225 ka, associated with handaxe and levallois artefacts. Their Neanderthal characteristics and their affinities with other European Middle Pleistocene teeth and the major sites of Atapuerca‐SH in Spain and Krapina in Croatia are explored. The Pontnewydd teeth are shown to have a marked simil...